論文

Kenji Suetsugu*, Satoshi Kakishima, and Yudai Okuyama. (2025). Seed dispersal by cockroaches and crickets, possibly attracted through yeasty scents, in the non-photosynthetic plant Balanophora tobiracola. Ecology. In Press.

奥山雄大　2026　カンアオイの遺伝子に刻まれた「虫を騙す香り」　岩波書店「科学」96号　pp. 65-70.

奥山雄大　2025　「擬態する花」を可能にする分子メカニズムと，その進化的起源　植物科学の最前線（BSJ-Review）vol. 16.

Hiroki Nishigaki*, Tetsuya K. Matsumoto, Yuko Miyazaki, Muneto Hirobe, Satoshi Kakishima, Yudai Okuyama, and Kenji Suetsugu*. (2025). Ethological evidence for sexually exploitative pollinator attraction by Arisaema serratum (Araceae). Journal of Plant Research, In Press.

Naoko Okui, Satoshi Kakishima, and Yudai Okuyama.* 2025. A unique pollination mode of Spuriomitella (H.Boissieu) R.A.Folk & Y.Okuyama observed in Japan’s Tohoku region. Bulletin of the National Museum of Nature and Science. Series B, Botany. 52: 1-9.

Jui-Tse Chang*, Tsung-Chen Lee, Anna K. Valchanova, Tomoko Okamoto, Pei-Chun Liao*, Goro Kokubugata, and Yudai Okuyama.* 2025. Match of fruit-scented cone volatile compositions with genetic boundary in Cycas revoluta and implications for fruit mimicry pollination. Plant Biology. 27: 725-739

Yudai Okuyama*, Kenji Fukushima, Satoshi Kakishima, Anna K. Valchanova, Kohei Takenaka Takano, Yasuko Ito-Inaba, Takeru Nakazato, and Atsushi J. Nagano. 2025. Convergent acquisition of disulfide-forming enzymes in malodorous flowers. Science. 388: 656-661.

奥山雄大　2025　花の香りの進化が駆動する植物の種分化　FFIジャーナル230巻1号　pp. 24-30.

奥山雄大　2024　チャルメルソウ−キノコバエ−コケ　思いがけず見つかった三者系　昆虫と自然 59: 15-18.

Anna K. Valchanova, Satoshi Kakishima, Yudai Okuyama.* 2024. Floral scent profiles and visits of Cordyla murina (Mycetophilidae) to Asarum curvistigma (Aristolochiaceae). Bulletin of the National Museum of Nature and Science. Series B, Botany. 50: 113-120.

Satoshi Kakishima*, Jun-ichi Ohno, Hoshiyama, K., Yamane, F., Sugie, Y., Yudai Okuyama.* 2023. Rediscovery of Arisaema suwoense Nakai in Yamaguchi Prefecture reveals morphological differences from A. izuense Nakai. Bulletin of the National Museum of Nature and Science. Series B, Botany. 49. 105-114.

奥山雄大　2023　チャルメルソウ・カンアオイ・テンナンショウ—「ハエ」を操り繁栄した日本の固有植物たち　岩波書店「科学」93号　pp. 340-344.

Ryan A. Folk, Gaynor, M. L., Engle-Wrye, N. J., O'Meara, B. C., Soltis, P. S., Soltis, D. E., R.P. Guralnick, S.A. Smith, C.J. Grady, Y., Okuyama, Y. 2023. Identifying climatic drivers of hybridization with a new ancestral niche reconstruction method. Systematic Biology 72: 856-873.

Yudai Okuyama*, Yuta Kato. 2022. Diversity of microscopic structures on the calyx of sect. Heterotropa (Aristolochiaceae). Bulletin of the National Museum of Nature and Science. Series B, Botany. 48. 133-139.

Yudai Okuyama*, Satoshi Kakishima. 2022. Possible adaptive and non-adaptive radiation in three plant genera in the Japanese Archipelago. Population Ecology. 64: 130-135.

Satoshi Kakishima, Masahiro Sueyoshi, Yudai Okuyama*. 2021. Floral visits of Cordyla murina (Mycetophilidae) and other dipterans to Asarum asaroides (Aristolochiaceae) and the possible role of mushroom-like scents. Bulletin of the National Museum of Nature and Science. Series B, Botany. 47: 227-236. DOI: 10.50826/bnmnsbot.47.4_227

Ryan A. Folk*, Rebecca L. Stubbs, Nicholas J. Engle-Wrye, Douglas E. Soltis, Yudai Okuyama*. 2021. Biogeography and habitat evolution of Saxifragaceae, with a revision of generic limits and a new tribal system. Taxon. 70: 263-285. DOI: 10.1002/tax.12450

Kenji Suetsugu*, Rikuo Sato, Satoshi Kakishima, Yudai Okuyama, Masahiro Sueyoshi. 2021. The sterile appendix of two sympatric Arisaema species lures each specific pollinator into deadly trap flowers. Ecology. doi:10.1002/ecy.3242

Satoshi Kakishima, Yudai Okuyama*. 2020. Further insights into the floral biology of Asarum tamaense (sect. Heterotropa, Aristolochiaceae). Bulletin of the National Museum of Nature and Science. Series B, Botany. 46: 129-143. doi: 10.50826/bnmnsbot.47.4_227

Yudai Okuyama*, Nana Goto, Atsushi Nagano, Masaki Yasugi, Goro Kokubugata, Hiroshi Kudo, Zhechen Qi, Takuro Ito, Satoshi Kakishima, Takashi Sugawara. 2020. Radiation history of Asian Asarum (sect. Heterotropa, Aristolochiaceae) resolved using a phylogenomic approach based on double-digested RAD-seq data. Annals of Botany. 126: 245-260.

Satoshi Kakishima, Masahiro Sueyoshi, Yudai Okuyama*. 2020. Floral visitors of critically endangered Arisaema cucullatum (Araceae) endemic to Kinki Region of Japan. Bulletin of the National Museum of Nature and Science. Series B, Botany. 46: 47-53.

Satoshi Kakishima, Yi-shuo Liang, Takuro Ito, T.-Y. Aleck Yang, Yudai Okuyama, Mitsuyasu Hasebe, Jin Murata, Jin Yoshimura*. 2019. Evolutionary origin of a periodical mass-flowering plant. Ecology and Evolution 9: 4373-4381.

Satoshi Kakishima, Yudai Okuyama*. 2018. Pollinator assemblages of Arisaema heterocephalum subsp. majus (Araceae), a critically endangered species endemic to Tokunoshima Island, Central Ryukyus. Bulletin of the National Museum of Nature and Science. Series B, Botany 44: 173-179.(333.6 KB)

Kiyotaka Hori*, Yudai Okuyama, Yasuyuki Watano, Noriaki Murakami. 2018. Recurrent hybridization without homoeologous chromosome paring in the Dryopteris varia complex (Dryopteridaceae). Chromosome Botany, 13: 9-24.

Yudai Okuyama*, Tomoko Okamoto, Jostein Kjærandsen, Makoto Kato. 2018. Bryophytes facilitate outcrossing of Mitella by functioning as larval food for pollinating fungus gnats. Ecology 99: 1890-1893.

Satoshi Kakishima, Yudai Okuyama*. 2018. Floral scent profiles and flower visitors in species of Asarum series Sakawanum (Aristolochiaceae). Bulletin of the National Museum of Nature and Science. Series B, Botany 44: 41–51(807.0 KB)

Yudai Okuyama*. 2016. Compartmentalized floral scent emission in two species of Mitella (Saxifragaceae). Bulletin of the National Museum of Nature and Science. Series B, Botany 42: 77-82.(414.3 KB)

Ryohei Terauchi*, Hiroyuki Kanzaki, Koki Fujisaki, Hiroki Takagi, Akira Abe, Kentaro Yoshida, Yudai Okuyama, Muluneh Tamiru, Hiromasa Saitoh. 2016. Whole genome sequencing approaches to understand Magnaporthe-rice interactions. Physiological and Molecular Plant Pathology. 95: 4-7.

Yudai Okuyama*. 2016. Mitella amamiana sp. nov., the first discovery of the genus Mitella (Saxifragaceae) in the central Ryukyus. Acta Phytotaxonomica et Geobotanica 67: 17-27.(4.3 MB)

Yudai Okuyama*. 2015. A consistent phenotyping of floral scent in Mitella (Saxifragaceae) using solid-phase microextraction. Bulletin of the National Museum of Nature and Science. Series B, Botany 41: 125-129.(226.0 KB)

奥山雄大*. 2015.日本産チャルメルソウ属およひ近縁種(ユキノシタ科)の自然史 Bunrui 15:109-123.(1.3 MB)

Tomoko Okamoto, Yudai Okuyama*, Ryutaro Goto, Masahiko Tokoro, Makoto Kato. 2015. Parallel chemical switches underlying pollinator isolation in Asian Mitella. Journal of Evolutionary Biology 28: 590-600.(737.9 KB)

Kenji Suetsugu*, Koji Tanaka, Yudai Okuyama, Tomohisa Yukawa. 2015. Potential pollinator of Vanda falcata (Orchidaceae): Theretra (Lepidoptera: Sphingidae) hawkmoths are visitors of the long spurred orchid. European Journal of Entomology, in press

Yudai Okuyama*. 2014. Repeated visitation by a sesarmid crab to male inflorescences of Piper sp. Bulletin of the National Museum of Nature and Science. Series B, Botany 40: 119-123.(4.1 MB)

Yudai Okuyama*, Mio Akashi. 2013. The genetic basis of flower-related phenotypic differences between closely related species of Asian Mitella (Saxifragaceae). Bulletin of the National Museum of Nature and Science. Series B, Botany 39: 131-136.(6.2 MB)

Stella Cesari, Gaëtan Thilliez, Cécile Ribot, Véronique Chalvon, Corinne Michel, Alain Jauneau, Susana Rivas, Ludovic Alaux, Hiroyuki Kanzaki, Yudai Okuyama, Jean-Benoit Morel, Elisabeth Fournier, Didier Tharreau, Ryohei Terauchi and Thomas Kroj*. 2013. The rice resistance protein pair RGA4/RGA5 recognizes the Magnaporthe oryzae effectors AVR-Pia and AVR1-CO39 by direct binding. The Plant Cell. 25: 1463-1481.

Yudai Okuyama*. 2012. Pollination by fungus gnats in Mitella formosana (Saxifragaceae). Bulletin of the National Museum of Nature and Science. Series B, Botany 38: 193-197.(1.7 MB)

Takayuki Mizuno*, Yudai Okuyama, Tsukasa Iwashina. 2012. Phenolic compounds from Iris rossii, and their chemotaxonomic and systematic significance. Biochemical Systematics and Ecology 44: 157-160.

Atsushi J. Nagano, Takashi Tsuchimatsu, Yudai Okuyama, Ikuko Hara-Nishimura*. 2012. Bimodal expression level polymorphisms in Arabidopsis thaliana. Plant Signaling and Behavior 7: 864-873.

Chisato Kobayashi*, Yudai Okuyama, Kazuhide Kawazoe, Makoto Kato. 2012. The evolutionary history of maternal plant-manipulation and larval feeding behaviours in attelabid weevils (Coleoptera; Curculionoidea). Molecular Phylogenetics and Evolution 318-330.

奥山雄大　2012　チャルメルソウ類を通して見る日本固有種の多様化　Bunrui 12: 31–39.

Yudai Okuyama*, Kunihiko Uno. 2012. The genetic characteristics of an endangered population of Mitella furusei var. subramosa (Saxifragaceae) from Shikoku, Japan. Bulletin of the National Museum of Nature and Science. Series B, Botany 38. 19-27.(1.5 MB)

Yudai Okuyama*, Akifumi S. Tanabe, Makoto Kato. 2012. Entangling ancient allotetraploidization in Asian Mitella: An integrated approach for multilocus combinations. Molecular Biology and Evolution 29: 429-439.

Ryohei Terauchi*, Kentaro Yoshida, Hiromasa Saitoh, Hiroyuki Kanzaki, Yudai Okuyama, Koki Fujisaki, Ayako Miya, Akira Abe, Muluneh Tamiru, Yukio Tosa. 2011. Studying genome-wide DNA polymorphisms to understand Magnaporthe-rice interactions. Australian Plant Pathology 40: 328-334.

Yudai Okuyama, Hiroyuki Kanzaki, Akira Abe, Kentaro Yoshida, Muluneh Tamiru, Hiromasa Saitoh, Takahiro Fujibe, Hideo Matsumura, Matt Shenton, Dominique Clark Galam, Jerwin Undan, Akiko Ito, Teruo Sone, Ryohei Terauchi*. 2011. A multi-faceted genomics approach allows the isolation of rice Pia-blast resistance gene consisting of two adjacent NBS-LRR protein genes. The Plant Journal 66: 467-479.

Makoto Shirakawa, Haruko Ueda, Tomoo Shimada, Yasuko Koumoto, Takashi L. Shimada, Maki Kondo, Taku Takahashi, Yudai Okuyama, Mikio Nishimura, Ikuko Hara-Nishimura*. 2010. Arabidopsis Qa-SNARE SYP2 proteins localized to different subcellular regions function redundantly in vacuolar protein sorting and plant development. The Plant Journal 64: 924-935.

Yudai Okuyama*. 2010. Which genus to study? In search of plant genera underrepresented or overrepresented in the research from the flora of Japan. Bulletin of the National Museum of Nature and Science. Series B, Botany 36:82-89.(206.1 KB)

Yudai Okuyama*, Makoto Kato. 2009. Unveiling cryptic species diversity of flowering plants: successful species circumscription of Asian Mitella using nuclear ribosomal DNA sequences. BMC Evolutionary Biology 9: 105(1.2 MB)

Makoto Kato*, Yasuyuki Kosaka, Atsushi Kawakita, Yudai Okuyama, Chisato Kobayashi, Thavy Phimminith, Daovorn Thongphan. 2008. Plant-pollinator interactions in tropical monsoon forests in Southeast Asia. American Journal of Botany 95: 1375-1394.

Yudai Okuyama*, Olle Pellmyr, Makoto Kato. 2008. Parallel floral adaptations to pollination by fungus gnats within the genus Mitella (Saxifragaceae). Molecular Phylogenetics and Evolution 46: 560-575

Yudai Okuyama*, Noriyuki Fujii, Michio Wakabayashi, Atsushi Kawakita, Manabu Ito, Mikio Watanabe, Noriaki Murakami, Makoto Kato. 2005. Nonuniform concerted evolution and chloroplast capture: heterogeneity of observed introgression patterns in three molecular data partition phylogenies of Asian Mitella (Saxifragaceae). Molecular Biology and Evolution 22: 285-296.(596.9 KB)

Makoto Kato*, Yudai Okuyama. 2004. Changes in the biodiversity of a deciduous forest ecosystem caused by an increase in the Sika deer population at Ashu, Japan. Contribution from the Biological Laboratory, Kyoto University 29: 437-448.

Yudai Okuyama*, Makoto Kato, Noriaki Murakami. 2004. Pollination by fungus gnats in four species of the genus Mitella (Saxifragaceae). Botanical Journal of the Linnean Society 144: 449-460.(561.9 KB)