論文

Ebihara, A., R. Koda and D. T. Vasques, 2025. Lectotypification of six fern taxa described by Tomitaro Makino based on specimen investigation at MAK, TI and TNS herbaria, Journal of Japanese Botany, 100: 40-51.
Lee, C. S., K.-S. Chung, S. Yang and A. Ebihara, 2024. Reticulate evolution in the Asplenium anogrammoides complex from Korea. Korean Journal of Plant Taxonomy, 54: 267-278.
Ebihara, A., N. Nakato and N. Inoue, 2024. A new record of the interspecific hybrid between Asplenium pekinense and A. varians (Aspleniaceae) in Japan. Bulletin of the National Museum of Nature and Science, Series B (Botany) , 50: 105-112.
Uemura, H., A. Ebihara, T. Hosoya, G. Kokubugata, T. Mizuno, Y. Murai, Y. Okuyama, N. Tanaka, N. Tanaka, C. Tsutsumi and T. Yukawa, 2024. A list of vascular plants in Tsukuba Botanical Garden 2023. Bulletin of the National Museum of Nature and Science, Series B (Botany), 50: 165-168. [Data paper]
Zhao, Y., E. Boucheron-Dubuisson, G. Rouhan, K. Iwatsuki, A. Ebihara, S. Hennequin and J.-Y. Dubuisson, 2025. Updated diversity and taxonomy of the fern genus Abrodictyum C. Presl (Hymenophyllaceae, Polypodiidae) in Asia, Australasia and Oceania. Botany Letters, 172: 63–86.
Suetsugu, K., H. Okada, S. K. Hirota, M. Yamasaki, R. Imaichi and A. Ebihara, 2025. Drastic mycorrhizal community shifts in Sceptridium ferns during the generation transition from fully mycoheterotrophic gametophytes to photosynthetic sporophytes. New Phytologist, 245: 1705-1717.
Vasques, D. T., A. Ebihara, A. Takano, H. Ikeda and A. Kawakita, 2024. History and importance of the fern herbarium collections in Japan, with focus on the University of Tokyo herbarium. Biodiversity Information Science and Standards, 8: e138439.
早川宗志・杉野孝雄・海老原淳, 2024. アミシダ（ヒメシダ科）の北限かつ東限を更新. 植物研究雑誌, 99: 268-270.
Yamamoto, K., T. Oka, A. Ebihara, S. Matsumoto, N. Murakami and K. Hori, 2024. Dryopteris ×makabensis (Dryopteridaceae), a new hybrid between Dryopteris caudipinna and D. tokyoensis. Acta Phytotaxonomica et Geobotanica, 75: 1-9.
Kuo, L.-Y., H.-J. Su, D. Koubinova, P.-J. Xie, C. Whitehouse, A. Ebihara and J. R. Grant, 2024. Organellar phylogenomics of Ophioglossaceae fern genera. Frontiers in Plant Science, 14: 1294716.
Boucheron-Dubuisson, E., Y. Zhao, G. Rouhan, A. Ebihara, K. Iwatsuki, D. Darnaedi and J.-Y. Dubuisson, 2024. The first record of Trichomanes s.s. (Hymenophyllaceae) in Asia and establishment of the new subgenus Asiatrichomanes. Phytotaxa 633: 215-229.
Ebihara, A., T. Fujiwara, M. Takamiya, M. Ito and T. Yahara, 2023. FernGreenList ver. 2.0: An updated checklist of wild fern and lycophyte species in Japan. Bulletin of National Museum of Nature and Science, Series B, 49: 97–104.
Naiki, A., T. Yamamoto, A. Hasegawa, T. Fujiwara, K. Yoneoka and A. Ebihara, 2023. Hymenophyllum pilosissimum C. Chr. (Hymenophyllaceae) on Iriomote Island, a new record for Japan. Acta Phytotaxonomica et Geobotanica, 74: 131–133.
Duibuisson, J.-Y., A. Nivart, E. Bidault, V. Deblauwe, V. Droissart, N. G. Kamdem, G. Rouhan, A. Ebihara and T. le Pechon, 2023. Diversity, taxonomy, and history of the tropical fern genus Didymoglossum Desv. (Hymenophyllaceae, Polypodiidae) in Africa. Journal of Systematics and Evolution, 62: 84–101.
Ebihara, A., S. Tagane, S. K. Hirota, Y. Suyama, N. Nakato and L.-Y. Kuo, 2023. Resurrection of Spicantopsis hancockii (Blechnaceae) as an endemic species to Taiwan: Reidentification of Spicantopsis in the Tokara Islands, Japan. Taiwania, 68: 148–154.
Li, Y., A. Ebihara, N. Nosova, Z.-Z. Tan and Y.-M. Cui, 2023. First fossil record of Trichomanes sensu lato (Hymenophyllaceae) from the mid-Cretaceous Kachin Amber, Myanmar. Life, 13: 1709.
Yoneoka, K., K. Hori, T. Kataoka, T. Fujiwara, A. Ebihara and N. Murakami, 2023. Hidden diversity of ferns: Haplopteris mediosora have survived as independent gametophytes in Japan. Acta Phytotaxonomica et Geobotanica, 74: 1-15.
Fujiwara, T., H. Liu, E. I. Meza-Torres, R. E. Morero, A. J. Vega, Z. Liang, A. Ebihara, I. J. Leitch and H. Schneideer, 2023. Evolution of genome space occupation in ferns: linking genome diversity and species richness. Annals of Botany, 131: 59–70.
Ebihara, A. and T. Nakasone. 2022. Hymenophyllum digitatum (Hymenophyllaceae) in Ishigaki-jima Island, a New Record for Japan. Journal of Japanese Botany, 97: 323-328.
Fujiwara, T., K. Yoneoka, Z. Liang, A. Ebihara, H. Schneider, N. Murakami and Y. Watano. 2022. Lepisorus rufofuscus sp. nov. (Polypodiaceae), a new fern species segregated from L. angustus Ching. Acta Phytotaxonomica et Geobotanica, 73: 171-182.
Vasques, D. T. and A. Ebihara. 2022. Phylogeny, morphology and distribution of the Hymenophyllum polyanthos (Sw.) Sw. species complex (Hymenophyllaceae, Polypodiidae) in Japan. Taiwania, 67: 524-538.
Lu, J.-M., X.-Y. Du, L.-Y. Kuo, A. Ebihara, L. R. Perrie, Z.-Y. Zuo, H. Shang, Y.-H. Chang and D.-Z.Li. 2022. Plastome phylogenomic analysis reveals evolutionary divergences of Polypodialessuborder Dennstaedtiineae. MBC Plant Biology, 22: 511.
Fujiwara, T., T. Egashira, J. S. Guiterrez-Ortega, K. Hori, A. Ebihara and Y. Watano. Establishment of an allotetraploid fern species, Lepisorus yamaokae Seriz., between two highly niche‐differentiated parental species. American Journal of Botany, 109: 1456-1471.
Dubuisson, J.-Y., E. Boucheron-Dubuisson, T. Le Pechon, P. Bausero, V. Droissart, V. Deblauwe, A. Ebihara, S. Hennequin and G. Rouhan. 2022. Diversity and taxonomy of the fern genus Vandenboschia Copel. (Hymenophyllaceae, Polypodiidae) in the Afro-Malagasy region and description of a new species. Botany Letters, 169: 268-283.
Chao, Y.-S., A. Ebihara, W.-L. Chiou, J.-M. Tsai, Y.-W. Huang and T. A. Ranker. 2022. Reticulate evolution in the Pteris fauriei group (Pteridaceae). Scientific Reports, 12: 9145.
Nitta, J. H., B. D. Mishler, W. Iwasaki and A. Ebihara. 2022. Spatial phylogenetics of Japanese ferns: Patterns, processes, and implications for conservation. American Journal of Botany, 109: 727-745.
Ebihara, A., Y.-S. Chao and H. Kanemitsu. 2022. The Distribution of Pteris wulaiensis (Pteridaceae) in Japan. Bulletin of National Museum of Nature and Science, Series B, 48: 53-59.
Nakagawa, H., H. Nagamasu, S. Nemoto, S. Fuse, A. Ebihara and K. Shutoh. 2022. Leymus komarovii (Triticeae, Poaceae) in Japan. Acta Phytotaxonomica et Geobotanica, 73: 151-157.Dubuisson, J.-Y., T. Le Pechoh, S. Hennequin, G. Rouhan, A. Salino, V. Deblauwe, V. Droissart, H. Tuomisto, S. Lehtonen and A. Ebihara. 2022. New insights into the diversity, taxonomy and history of the fern genus Trichomanes (Hymenophyllaceae, Polypodiidae), with a focus on Africa and the western Indian Ocean. Botanical Journal of the Linnean Society, 198: 215-239.
Dubuisson, J.-Y., T. Le Pechoh, S. Hennequin, G. Rouhan, A. Salino, V. Deblauwe, V. Droissart, H. Tuomisto, S. Lehtonen and A. Ebihara. 2022. New insights into the diversity, taxonomy and history of the fern genus Trichomanes (Hymenophyllaceae, Polypodiidae), with a focus on Africa and the western Indian Ocean. Botanical Journal of the Linnean Society, 198: 215-239.
大森威宏・黒沢高秀・志賀隆・薄葉満・根本秀一・吉井広始・海老原淳・田中徳久・天野誠. 2022. 尾瀬の維管束植物目録の見直し. 低温科学, 80: 199-223.
大森威宏・黒沢高秀・志賀隆・薄葉満・根本秀一・吉井広始・海老原淳・田中徳久・天野誠. 2022. 尾瀬産維管束植物相とその再検討. 低温科学, 80: 175-197.中藤成実・海老原淳. 2021. ヘラシダ（メシダ科）の二倍体・四倍体・六倍体は胞子のサイズと表面模様で識別できる. 植物地理・分類研究, 69: 233-248.
中藤成実・海老原淳. 2021. ヘラシダ（メシダ科）の二倍体・四倍体・六倍体は胞子のサイズと表面模様で識別できる. 植物地理・分類研究, 69: 233-248.
Ebihara, A., N. Nakato, Y.-H. Chang and L.-Y. Kuo, 2021. Coryphopteris castanea (Thelypteridaceae), a polymorphic diploid species distributed in Taiwan and Japan. Bulletin of National Museum of Nature and Science, Series B, 47: 71-77.(1.3 MB)
Iwatsuki, K., M. Kato and A. Ebihara, 2021. A new filmy fern, Hymenophyllum seramense (Hymenophyllaceae), from Seram Island, Indonesia. Journal of Japanese Botany, 96: 229-232.
Fujiwara, T., J. Ogiso, S. Ishii, K. Togo, N. Nakato, S. Serizawa, Y.-S. Chao, H.-T. Im, A. Ebihara and Y. Watano, 2021. Species delimitation in the Phegopteris decursivepinnata polyploid species complex (Thelypteridaceae). Acta Phytotaxonomica et Geobotanica, 72: 205-226.
Chao, Y.-S., W.-L. Chiou, A. Ebihara, T.-C. Hsu, Y.-H. Chang and C.-Y. Lin, 2021. Taxonomic and nomenclatural novelties in the Pteris fauriei group (Pteridaceae). Taiwania, 66: 307-316.
Wu, Y.-H., C.-Y. Sun, A. Ebihara, N. T. Lu, G. Rouhan and L.-Y. Kuo, 2021. Two new species in the fern genus Lomariopsis (Lomariopsidaceae) from East Asia. PhytoKeys, 187: 161-176.
Zhang, M.-H., R. Wei, Q.-P. Xiang, A. Ebihara and X.-C. Zhang, 2021. Integrative taxonomy of the Selaginella helvetica group based on morphological, molecular and ecological data. Taxon, 70: 1163-1187.
Kuo, L.-Y., S. K. Tang, T.-T. kao, A. Ebihara, S. Faqcett, M.-C. Hsiao, W. Shinohara and B. Dauphin, 2021. A dormant resource for genome size estimation in ferns: C‐value inference of the Ophioglossaceae using herbarium specimen spores. Applications in Plant Sciences, 9: e11452.Zhao, T., Z.-Y. Zuo, A. Ebihara, N. Nakato, A. Soejima, D.-Z. Li, J. Wen and J.-M. Lu, 2021. Adiantum japonicum, a new species of the Adiantum pedatum complex (Pteridaceae) from Japan. Phytotaxa, 525: 1-14.
Zhao, T., Z.-Y. Zuo, A. Ebihara, N. Nakato, A. Soejima, D.-Z. Li, J. Wen and J.-M. Lu, 2021. Adiantum japonicum, a new species of the Adiantum pedatum complex (Pteridaceae) from Japan. Phytotaxa, 525: 1-14.
Chen, C.-W,. A. Ebihara, K.-Y. Cheng, T.-C. Hsu, Y.-M. Huang, V. D. Dang, H. T. Luu, V. S. Le and C.-W. Li, 2021. Studies of Vietnamese pteridophyte flora 1. Systematic Botany, 46: 573-581.
杉田典正・海老原淳・細矢剛・神保宇嗣・中江雅典・遊川知久, 2021. 日本の絶滅危惧生物標本の所在把握と保全への活用. 保全生態学研究, 26: 229-247.
Ebihara, A., N. Nakato, T. Ohi-Toma & K. Iwatsuki, 2020. Thelypteris sylva-nipponica, a new allotetraploid species in the Parathelypteris group (Thelypteridaceae). Phytotaxa, 477: 229-242.
Nitta, J. H., A. Ebihara & A. R. Smith, 2020. A taxonomic and molecular survey of the pteridophytes of the Nectandra Cloud Forest Reserve, Costa Rica. PLoS ONE, 15(11): e0241231.
Nakato, N., A. Ebihara, M. Watanabe and C. Tsutsumi, 2020, New cytotaxonomic records on threatened fern species in Japan. Bulletin of the National Museum of Nature and Science, Series B, 46: 17–27.(1.2 MB)
Cámara-Leret, R., D. G. Frodin,F. Adema, C. Anderson, M. S. Appelhans, G. Argent, S. A. Guerrero, P. Ashton, W. J. Baker, A. S. Barfod, D. Barrington, R. Borosova, G. L. C. Bramley, M. Briggs, S. Buerki, D. Cahen, M. W. Callmander, M. Cheek, C.-W. Chen, B. J. Conn, M. J. E. Coode, I. D. S. Dawson, J. Dransfield, C. Drinkell, B. Duyfjes, A. Ebihara, Z. Ezedin, L.-F. Fu, O. Gideon, D. Girmansyah, R. Govaerts, H. Fortune-Hopkins, G. Hassemer, A. Hay, C. D. Heatubun, D. J. N. Hind, P. Hoch, P. Homot, P. Hovenkamp, M. Hughes, M. Jebb, L. Jennings, T. Jimbo, M. Kessler, R. Kiew, S. Knapp, P. Lamei, M. Lehnert, G. P. Lewis, H. P. Linder, S. Lindsay, Y. W. Low, E. Lucas, J. P. Mancera, A. K. Monro, A. Moore, D. J. Middleton, H. Nagamasu, M. F. Newman, E. N. Lughadha, P. H. A. Melo, D. J. Ohlsen, C. M. Pannell, B. Parris, L. Pearce, D. S. Penneys, L. R. Perrie, P. Petoe, A. D. Poulsen, G. T. Prance, J. P. Quakenbush, N. Raes, M. Rodda, Z. S. Rogers, A. Schuiteman, P. Schwartsburd, R. W. Scotland, M. P. Simmons, D. A. Simpson, P. Stevens, M. Sundue, W. Testo, A. Trias-Blasi, I. Turner, T. Utteridge, L. Walsingham, B. L. Webber, R. Wei, G. D. Weiblen, M. Weigend, P. Weston, W. de Wilde, P. Wilkie, C. M. Wilmot-Dear, H. P. Wilson, J. R. I. Wood, L.-B. Zhang & P. C. van Welzen, 2020. New Guinea has the world’s richest island flora. Nature 584: 579-583.
Sugita, N., A. Ebihara, T. Hosoya, U. Jinbo, S. Kaneko, T. Kurosawa, M. Nakae & T. Yukawa, 2020. Non‑destructive DNA extraction from herbarium specimens: a method particularly suitable for plants with small and fragile leaves. Journal of Plant Research, 133: 133-141.
Kuo, L.-Y., Y.-H. Chang, Y.-H. Huang, W. Testo, A. Ebihara, G. Rouhan, L. Quintanilla, J. E. Watkins Jr., Y.-M. Huang and F.-W. Li, 2019. A global phylogeny of Stegnogramma ferns (Thelypteridaceae): generic and sectional revision, historical biogeography and evolution of leaf architecture. Cladistics: 36: 164-183.
Ebihara, A. and J. H. Nitta, 2019. An update and reassessment of fern and lycophyte diversity data in the Japanese Archipelago. Journal of Plant Research 132: 723–738.
Vasques, D. T., A. Ebihara, R. Y. Hirai, J. Prado and M. Ito, 2019. Phylogeny of Hymenophyllum subg. Mecodium (Hymenophyllaceae), with special focus on the diversity of the Hymenophyllum polyanthos species complex. Plant Systematics and Evolution, 305: 811-825.
Liu, H.-M., L. Ekrt, P. Koutecky, J. Pellicer, O. Hidalgo, J. Marquardt, F. Pustahija, A. Ebihara, S. Siljak‐Yakovlev, M. Gibby, I. Leitch and H. Schneider, 2019. Polyploidy does not control all: Lineage‐specific average chromosome length constrains genome size evolution in ferns. Journal of Systematics and Evolution, 57: 418-430.
Ogura-Tsujita, Y., K. Yamamoto, Y. Hirayama, A. Ebihara, N. Morita and R. Imaichi, 2019. Fern gametophytes of Angiopteris lygodiifolia and Osmunda japonica harbor diverse Mucoromycotina fungi. Journal of Plant Research, 132: 581-588.
Ebihara, A., J. H. Nitta, Y. Matsumoto, Y. Fukazawa, M. Kurihara, H. Yokote, K. Sakuma, O. Azakami, Y. Hirayama and R. Imaichi, 2019. Growth dynamics of independent gametophytes of Pleurosoriopsis makinoi (Polypodiaceae). Bulletin of the National Museum of Nature and Science, Series B, 45: 77-86.(1.3 MB)
Ebihara, A., J. H. Nitta, Y. Matsumoto, Y. Fukazawa, M. Kurihara, H. Yokote, K. Sakuma, O. Azakami, Y. Hirayama and R. Imaichi, 2019. Growth dynamics of independent gametophytes of Pleurosoriopsis makinoi (Polypodiaceae). Bulletin of the National Museum of Nature and Science, Series B, 45: 77-86.(1.3 MB)
Iwatsuki, K., A. Ebihara and M. Kato, 2019. Taxonomic studies of pteridophytes of Ambon and Seram (Moluccas) collected on Indonesian-Japanese botanical expeditions 1983-1986. XIII. Hymenophyllaceae. PhytoKeys, 119: 107-115.
Hori, K., M. Watanabe, A. Ebihara, I. Yamazumi, M. Takamiya and N. Murakami, 2019. Genome constitution of the Dryopteris atrata complex (Dryopteridaceae). Cytlogia, 84: 135-141.
Ebihara, A., N. Nakato, L.-Y. Kuo, H. Miyazaki and S. Serizawa, 2019. Allopolyploid origin and distribution range of Acystopteris taiwaniana (Cystopteridaceae: Polypodiales). Acta Phytotaxonomica et Geobotanica 70: 19-28.
Lu, N. T., A. Ebihara, H. He, L. Zhang, Z,-M. Zhou, R. Knapp, P. Kamau, D. Lorence, X.-F. Gao and L.-B. Zhang, 2019. A plastid phylogeny of the fern genus Arachniodes (Dryopteridaceae). Molecular Phylogenetics and Evolution 133: 214-235.
上野勝典・上野由貴枝・海老原淳, 2019. Dennstaedtia punctilobula（コバノイシカグマ科）が長野県に帰化. Journal of Japanese Botany 94: 58-60.
Hori, K., A. Ebihara and A. Z. Moe, 2019. New records of Pteridophytes from Myanmar (1). Journal of Japanese Botany 94: 35-38.
海老原淳, 2019. 『日本産シダ植物標準図鑑』の分布図ができるまで. 東北植物研究 20： 1-11.
海老原淳・森田秀一・山室一樹, 2018. 徳之島で発見された日本新産種ムシャシダ（メシダ科）. Journal of Japanese Botany 93: 404-406.
Hori, K., X.-L. Zhou, W. Shao, Y.-H. Yan, A. Ebihara, R.-X. Wang, H. Ishikawa, Y. Watano and N. Murakami, 2018. Hybridization of the Dryopteris erythrosora complex (Dryopteridaceae, Polypodiidae) in Japan and adjacent areas. Hikobia 17: 299-313.
Kuo, L.-Y., A. Ebihara, T.-C. Hsu, G. Rouhan, Y.-M. Huang, C.-N. Wang, W.-L. Chiou and S. Matsumoto, 2018. Infrageneric revision of the fern genus Deparia (Athyriaceae, Aspleniineae, Polypodiales). Systematic Botany 43: 645-655.
Chang, Y., A. Ebihara, S. Lu, H. Liu and H. Schneider, 2018. Integrated taxonomy of the Asplenium normale complex (Aspleniaceae) in China and adjacent areas. Journal of Plant Research 131: 573-587.
Hori, K., A. Ebihara and N. Murakami, 2018. Revised classification of the species within the Dryopteris varia complex (Dryopteridaceae) in Japan. Acta Phytotaxonomica et Geobotanica 69: 77-108.
Nakato, N. and A. Ebihara, 2018. Chromosome numbers of eleven ferns in Japan (Athyriaceae, Dryopteridaceae and Tectariaceae). Bulletin of the National Museum of Nature and Science, Series B, 44: 23-30.(10.7 MB)
Kuo, L.-Y., A. Ebihara, M. Kato, G. Rouhan, T. A. Ranker, C.-N. Wang and W.-L. Chiou, 2018. Morphological characterization of infra-generic lineages in Deparia (Athyriaceae: Polypodiales). Cladistics 34: 78-92.
Wei, R, A. Ebihara, Y.-M. Zhu, C.-F. Zhao, S. Hennequin and X.-C. Zhang, 2018. A total-evidence phylogeny of the lady fern genus Athyrium Roth (Athyriaceae) with a new infrageneric classification. Molecular Phylogenetics and Evolution 119: 25-36.Ranil, R. H. G., D. K. N. G. Pushpakumara, D. S. A. Wjiesundara, P. D. Bostock, A. Ebihara and C. R. Fraser-Jenkins, 2017. Diversity and distributional ecology of tree ferns of Sri Lanka: A step towards conservation of a unique gene pool. Ceylon Journal of Science 46 (Special Issue) 2017: 127-135.
Ranil, R. H. G., D. K. N. G. Pushpakumara, D. S. A. Wjiesundara, P. D. Bostock, A. Ebihara and C. R. Fraser-Jenkins, 2017. Diversity and distributional ecology of tree ferns of Sri Lanka: A step towards conservation of a unique gene pool. Ceylon Journal of Science 46 (Special Issue) 2017: 127-135.
Chao, Y.-S., A. Ebihara, W.-L. Chiou, Y.-M. Huang, 2017. Pteris latipinna sp. nov. (Pteridaceae), a new species segregated from Pteris fauriei. PhytoKeys, 85: 95-108.
Dubuisson, J.-Y., L. Bauret, A. grall, B. Senterre, A.H. Said, K. Pynee, A. Ebihara and S. Hennequin, 2017. Taxonomic study of the fern genera Abrodictyum C.Presl and Trichomanes L. (Hymenophyllaceae, Polypodiidae) in the western Indian Ocean, and description of a new Abrodictyum species for Madagascar. Phytotaxa, 327: 201-222.
Del Rio, C., S. Hennequin, G. Rouhan, A. Ebihara, P.P. Lowry II, J.-Y. Dubuisson and M. Gaudeul, 2017. Origins of the fern genus Hymenophyllum (Hymenophyllaceae) in New Caledonia: Multiple independent colonizations from surrounding territories and limited in situ diversification. Taxon, 66: 1041-1064.
Kuo, L.-Y., C.-W. Chen, W. Shinohara, A. Ebihara, H. Kudoh, H. Sato, Y.-M. Huang and W.-L. Chiou, 2017. Not only in the temperate zone: independent gametophytes of two vittarioid ferns (Pteridaceae, Polypodiales) in East Asian subtropics. Journal of Plant Research, 130: 255-262.
Schneider, S., H.-M. Liu, Y.-F. Chang1 D. Ohlsen, L.R. Perrie, L. Shepherd, M. Kessler, D.N. Karger, S. Hennequin, J. Marquardt, S.Russell, S. Ansell, N.T. Lu, P. Kamau, J. Loriga, L. Regalad, J. Heinrichs, A. Ebihara, A.R. Smith and M. Gibby, 2017. Neo- and Paleopolyploidy contribute to the species diversity of Asplenium—the most species-rich genus of ferns. Journal of Systematics and Evolution, 55: 353-364.
Vasques, D.T., A. Ebihara and M. Ito, 2017. The felt fern genus Pyrrosia Mirbel (Polypodiaceae): a new subgeneric classification with a molecular phylogenic analysis based on three plastid markers. Acta Phytotaxonomica et Geobotanica, 68: 65-82.
Ebihara, A., N. Nakato and T. Jaruwattanaphan, 2017. A new taxonomic treatment for the apogamous counterpart of Pteris terminalis (Pteridaceae). Phytotaxa 314: 73-80.
Ebihara, A., S. Matsumoto, J. Mazumdar and K. Yamamoto, 2017. Updates of taxonomic treatments for ferns of Japan 2. Athyrium and Cyrtomium. Bulletin of the National Museum of Nature and Science, Series B, 43: 19-25.(1.0 MB)
Chen, C.-W., S. Lindsay, L.-Y. Kuo, C. R. Fraser-Jenkins, A Ebihara, H. T. Luu, C. W. Park, Y.-S. Chao, Y.-M. Huang and W.-L. Chiou, 2017. A systematic study of East Asian vittarioid ferns (Pteridaceae: Vittarioideae). Botanical Journal of the Linnean Society 183: 545-560.
Kuo, L.-Y., C.-W. Chen, W. Shinohara, A. Ebihara, H. Kudoh, H. Sato, Y.-M. Huang, W.-L. Chiou, 2017. Not only in the temperate zone: independent gametophytes of two vittarioid ferns (Pteridaceae, Polypodiales) in East Asian subtropics. Journal of Plant Research 130: 255-262.
Hori, K., L.-Y. Kuo, W.-L. Chiou, A. Ebihara and N. Murakami, 2017. Geographical distribution of sexual and apogamous types of Dryopteris formosana and Dryopteris varia (Dryopteridaceae) in Taiwan. Acta Phytotaxonomica et Geobotanica 68: 23-32.
Ebihara, A., N. Nakato, V. B. Amoroso, A. Hidayat and L.-Y. Kuo, 2016. Monachosorum arakii Tagawa (Dennstaedtiaceae) is a relict "international" hybrid: A reassessment of the Monachosorum species. Systematic Botany 41: 586-595.
Kuo, L.-Y., A. Ebihara, W. Shinohara, G. Rouhan, K. R. Wood., C.-N. Wang and W.-L. Chiou, 2016. Historical biogeography of the fern genus Deparia (Athyriaceae) and its relation with polyploidy. Molecular Phylogenetics and Evolution 104: 123–134.
Dubuisson, J.-Y., L. Bauret, A. Grall, T. Li, A. Ebihara and S. Hennequin, 2016. Discussion on the taxonomy of African fern Abrodictyum rigidum (Sw.) Ebihara & Dubuisson and description of two new Abrodictyum C.Presl species (Hymenophyllaceae, Polypodiidae) for the Afro-Malagasy region. Phytotaxa 284: 151-168.
The Pteridophyte Phylogeny Group, 2016. A community-derived classification for extant lycophytes and ferns. Journal of Systematics and Evolution, 54: 563-606.
Imai, R., Y. Tsuda, S. Matsumoto, A. Ebihara and Y. Watano, 2016. The relationship between mating system and genetic diversity in diploid sexual populations of Cyrtomium falcatum in Japan. Plos One 11(10): e0163683.
Wang, F.-H., J.-M. Lu, J. Wen, A. Ebihara and D.-Z. Li, 2016. Applying DNA barcodes to identify closely related species of ferns: a case study of the Chinese Adiantum (Pteridaceae). Plos One 11(9): e0160611.
Nakato, N. and A. Ebihara, 2016. Chromosome numbers of 18 ferns in Japan: toward completion of chromosome information in Japanese ferns. Bulletin of the National Museum of Nature and Science, Series B, 42: 25-40.(6.4 MB)
Ebihara, A., N. Nakato and S. Matsumoto, 2015. Updates of taxonomic treatments for ferns of Japan 1. Botrychium, Osmolindsaea and Pteris. Bulletin of the National Museum of Nature and Science, Series B, 41: 15-24.(554.9 KB)
Zhang, L., C.J. Rothfels, A. Ebihara, E. Schuettpelz, T.L. Pichon, P. Kamau, H. He, X.-M. Zhou, J. Prado, A. Field, G. Yatskievych, X.-F. Gao and L.-B. Zhang, 2015. A global plastid phylogeny of the brake fern genus Pteris (Pteridaceae) and related genera in the Pteridoideae. Cladistics, 31: 406-423.
Bauret, L., A. Grall, B. Senterre, G. Rouhan, S. Hennequin, A. Ebihara and J.-Y. Dubuisson, 2015. New circumscription of Trichomanes cupressoides Desvaux (Hymenophyllaceae), an endemic filmy fern from the Seychelles (Indian Ocean), and new insights into the genus Abrodictyum C.Presl in the western Indian Ocean. Phytotaxa 202: 1-14.
Hori., K., A. Ebihara, N. Nakato and N. Murakami, 2015. Dryopteris protobissetiana (Dryopteridaceae), a new diploid sexual species of the Dryopteris varia complex (subg. Erythrovariae, sect. Variae) from Yakushima, Kagoshima, Japan. Acta Phyotaxonomica et Geobotanica, 66: 47-57.
Hori, K., M. Matsumoto, A. Ebihara, N. Nakato and N. Murakami, 2015. Geographical distribution of sexual and apogamous types of Dryopteris chinensis (Dryopteridaceae) in Japan. Acta Phyotaxonomica et Geobotanica, 66: 35-45.
Ogura-Tsujita, Y., Y. Hirayama, A. Sakoda, A. Suzuki, A. Ebihara, N. Morita and R, Imaichi, 2015. Arbuscular mycorrhizal colonization in field-collected terrestrial cordate gametophytes of pre-polypod leptosporangiate ferns (Osmundaceae, Gleicheniaceae, Plagiogyriaceae, Cyatheaceae). Mycorrhiza, 26: 87-97.
Ebihara, A., N. Nakato, Y. Saito, T. Oka and T. Minamitani, 2014. New records of Asplenium varians (Aspleniaceae) and two new hybrids in Japan. Acta Phytotax. Geobot. 65: 53-65.
Ebihara, A., N. Nakato, S. Matsumoto, Y.-S. Chao and L.-Y. Kuo, 2014. Cytotaxonomic studies on thirteen ferns of Taiwan. Bull. Nat. Mus. Nat. Sci. B 40: 19-28.(5.3 MB)
Chen, C.-W., A. Ebihara, W.-L. Chiou and C.-W. Li, 2014. Haplopteris yakushimensis (Pteridaceae, Vittarioideae), a new species from Yakushima Island, Japan. Phytotaxa 156: 229-234.
Hori, K., A. Tono, K. Fujimoto, J. Kato, A. Ebihara, Y. Watano and N. Murakami, 2014. Reticulate evolution in the apogamous Dryopteris varia complex (Dryopteridaceae, subg. Erythrovariae, sect. Variae) and its related sexual species in Japan. Journal of Plant Research 127: 661–684.
Chao, Y.-S., A. Ebihara, Y.-H. Chang, R.-H. Jiang, L. T. Ngan and W.-L. Chiou, 2013. New distribution of Pteris kawabatae Sa. Kurata and re-circumscription of Pteris arisanensis Tagawa. Taiwania 58: 305-310.
Dubuisson, J.-Y., S. Bary, A. Ebihara, E. Carnero-Diaz, E. Boucheron-Dubuisson and S. Hennequin, 2013. Epiphytism, anatomy and regressive evolution in trichomanoid filmy ferns (Hymenophyllaceae). Bot. J. Linn. Soc. 173: 573-593.
Dubuisson, J.-Y., G. Rouhan, A. Gralla, S. Hennequin, B. Senterre, K. Pynee and A. Ebihara, 2013. New insights into the systematics and evolution of the filmy fern genus Crepidomanes (Hymenophyllaceae) in the Mascarene Archipelago with a focus on dwarf species. Acta Bot. Gallica 160: 173-194.
Ebihara, A., A. Yamaoka, N. Mizukami, A. Sakoda, J. H. Nitta and R. Imaichi, 2013. A survey of the fern gametophyte flora of Japan: Frequent independent occurrences of noncordiform gametophytes. Amer. J. Bot. 100: 735-743.
Ogura-Tsujita, Y., A. Sakoda, A. Ebihara, T. Yukawa and R. Imaichi, 2013. Arbuscular mycorrhiza formation in cordate gametophytes of two ferns, Angiopteris lygodiifolia and Osmunda japonica. J. Plant Res. 126: 41-50.
Ebihara, A. and N. Nakato, 2013. Distribution of the Thelypteris japonica Complex (Thelypteridaceae) in Japan. Bull. Natn. Mus. Nat. Sci. B, 39: 61-85.(1.1 MB)
Zhang, L.-B., L. Zhang, S.-Y. Dong, E. B. Sessa, X.-F. Gao and A. Ebihara, 2012. Molecular circumscription and major evolutionary lineages of the fern genus Dryopteris (Dryopteridaceae). BMC Evol. Biol. 12: 180.
Ebihara, A., S. Matsumoto and M. Kato, 2012. Origin of Dryopteris shibipedis (Dryopteridaceae), a fern species extinct in the wild. J. Plant Res. 125: 499-505.
Ebihara, A., C. R. Fraser-Jenkins, B. S. Parris, X.-C. Zhang, Y.-H. Yang, W.-L. Chiou, H.-M. Chang, S. Lindsay, D. Middleton, M. Kato, T. N. Praptosuwiryo, V. B. Amoroso, J. F. Barcelona, R. H. G. Ranil, C.-H. Park, N. Murakami and A. Hoya, 2012. Rare and threatened pteridophytes of Asia 1. An enumeration of narrowly distributed taxa. Bull. Natn. Mus. Nat. Sci. B, 38: 93-119.(490.4 KB)
Nitta, J. H., A. Ebihara and M. Ito, 2011. Reticulate evolution in the Crepidomanes minutum species complex. Amer. J. Bot. 98: 1782-1800.
Li, F.-W., L.-Y. Kuo, C. J. Rothfels, A. Ebihara, W.-L. Chiou, M. D. Windham and K. M. Pryer, 2011. rbcL and matK earn two thumbs up as the core DNA barcode for ferns. PLoS ONE 6: e26597.
Lin, S.-J., A. Ebihara, D. Wang and K. Iwatsuki, 2011. Systematic studies of Polystichum (Dryopteridaceae) in Japan (I): P. fibrillosopaleaceum var. marginale is a diploid hybrid between P. fibrillosopaleaceum and P. igaense. Amer. Fern J. 101: 295-306.
Ebihara, A, 2011. rbcL phylogeny of Japanese pteridophyte flora and implications on infrafamilial systematics. Bull. Natn. Mus. Nat. Sci. B, 37: 63-74.(948.9 KB)
Nakato, N. and A. Ebihara, 2011. Chromosome number of Microlepia hookeriana (Dennstaedtiaceae) and chromosome number evolution in the genus Microlepia. Bull. Natn. Mus. Nat. Sci. B, 37: 75-78.(333.9 KB)
Dubuisson, J.-Y., S. Hennequin, S. Bary, A. Ebihara and E. Boucheron-Dubuisson, 2011. Anatomical diversity and regressive evolution in trichomanoid filmy ferns (Hymenophyllaceae): A phylogenetic approach. Comptes Rendus Biologies 334: 880-895.
Lin, S.-J., T. Suzuki, A. Ebihara, H. Lu, J. Liu, S. Wu and K. Iwatsuki, 2011 A new record of Polystichum otomasui Sa.Kurata (Dryopteridaceae) from Fujian Province, China. J. Jpn. Bot. 86: 115-119.
Ebihara, A., J. H. Nitta and M. Ito, 2010. Molecular species identification with rich floristic sampling: DNA barcoding the pteridophyte flora of Japan. PLoS ONE 5(12): e15136.
Yamamoto, K., N. Murakami and A. Ebihara, 2010. The distribution of Dryopteris caudipinna (Dryopteridaceae), a sexually reproducing counterpart of the apogamous D. erythrosora, in Japan. Acta Phytotax. Geobot. 61: 109-114.
Hennequin, S., A. Ebihara, J.-Y. Dubuisson and H. Schneider, 2010. Chromosome number evolution of Hymenophyllum (Hymenophyllaceae), with special reference to the subgenus Hymenophyllum. Molec. Phylogen. Evol. 55: 47-59.
Ebihara, A., J. H. Nitta and K. Iwatsuki, 2010. The Hymenophyllaceae of the Pacific area. 2. Hymenophyllum (excluding subgen. Hymenophyllum). Bull. Natn. Mus. Nat. Sci., B, 36: 43-59.(2.9 MB)
Ebihara, A., J. H. Nitta, D. Lorence & J.-Y. Dubuisson, 2009. New records of Polyphlebium borbonicum, an African filmy fern, in the New World and Polynesia. Amer. Fern J. 99: 200-206.
Ebihara, A., S. Matsumoto & M. Ito, 2009. Hybridization involving independent gametophytes in the Vandenboschia radicans complex (Hymenophyllaceae): a new perspective on the distribution of fern hybrids. Molec. Ecol. 18: 4904-4911.
Ebihara, A., S. Matsumoto & M. Ito, 2009. Taxonomy of the reticulate Vandenboschia radicans complex (Hymenophyllaceae) in Japan. Acta Phytotax. Geobot. 60: 26-40.
Ebihara, A, 2009. Notes on the Distribution of the Vandenboschia radicans Complex (Hymenophyllaceae) in Japan. Bull. Natn. Mus. Nat. Sci., B, 35: 71-89.(638.2 KB)
Ebihara, A, 2008. Hymenophyllaceae collected in the Mt. Kinabalu Area in 2000 and 2007. Mem. Natn. Mus. Nat. Sci. 45: 105-110.
Ebihara, A., D. R. Farrar & M. Ito. 2008. The sporophyte-less filmy fern of eastern North America Trichomanes intricatum (Hymenophyllaceae) has the chloroplast genome of an Asian species. Amer. J. Bot. 95: 1645-1651.
Hennequin, S., E. Schuettpelz, K. M. Pryer, A. Ebihara & J.-Y. Dubuisson, 2008. Divergence times and the evolution of epiphytism in filmy ferns (Hymenophyllaceae) revisited. Int. J. Plant Sci. 169: 1278-1287.
Ebihara, A. & K. Miyashita, 2008. Type Collection in the Herbarium of National Museum of Nature and Science (TNS). Ericales, Primulales and Lamiales. Bull. Natn. Mus. Nat. Sci.,B, 33: 103-111.
Ebihara, A. & K. Miyashita, 2008. Type Collection in the Herbarium of National Museum of Nature and Science (TNS). Salicales, Fagales, Urticales, Proteales, Santalales, Polygonales and Centrospermae. Bull. Natn. Mus. Nat. Sci.,B, 33: 23-29.
Ebihara, A., K. Iwatsuki, M. Ito, S. Hennequin and J.-Y. Dubuisson, 2007. A global molecular phylogeny of the fern genus Trichomanes (Hymenophyllaceae) with special reference to stem anatomy. Botanical Journal of the Linnean Society, 155: 1-27.
Ebihara, A., K. Iwatsuki, M. Ito & J.-Y. Dubuisson, 2007. Systematics of Trichomanes (Hymenophyllaceae: Pteridophyta), progress and future interests. Fern Gazette, 18: 53-58.
Ebihara, A. & K. Iwatsuki, 2007. The Hymenophyllaceae of the Pacific Area 1. Hymenophyllum subgen. Hymenophyllum. Bull. Natn. Mus. Nat. Sci.,B, 33: 55-68.(2.9 MB)
Ebihara A., J.-Y. Dubuisson, K. Iwatsuki, S. Hennequin and M. Ito. 2006. A taxonomic revision of Hymenophyllaceae. Blumea 51: 221-280.
Hennequin S., A. Ebihara, M. Ito, K. Iwatsuki and J.-Y. Dubuisson. 2006. New insights into the phylogeny of the genus Hymenophyllum s.l. (Hymenophyllaceae): Revealing the polyphyly of Mecodium. Syst. Bot. 31: 271-284.
Hennequin S., A. Ebihara, M. Ito, K. Iwatsuki and J.-Y. Dubuisson. 2006. Phylogenetic systematics and evolution of the genus Hymenophyllum s.l. (Hymenophyllaceae: Pteridophyta). - Proceedings of the Ferns for the 21st Century Conference, Edinburgh 2004. Fern Gazette 17: 247-257.
Lin S.-J., A. Ebihara and K. Iwatsuki. 2006. Cytological observations on Lindsaea trichomanoides (Lindsaeaceae), a fern species from New Zealand. J. Jpn. Bot. 81: 103-106.
Ebihara A., H. Ishikawa, S. Matsumoto, S.-J. Lin, K. Iwatsuki, M. Takamiya, Y. Watano and M. Ito. 2005. Nuclear DNA, chloroplast DNA, and ploidy analysis clarified biological complexity of the Vandenboschia radicans complex (Hymenophyllaceae) in Japan and adjacent areas. Amer. J. Bot. 92: 1535-1547.
Ebihara A., S. Hennequin, K. Iwatsuki, P.D. Bostock, S. Matsumoto, R. Jaman, J.-Y. Dubuisson and M. Ito. 2004. Polyphyletic origin of Microtrichomanes (Prantl) Copel. (Hymenophyllaceae), with a revision of the species. Taxon 53: 935-948.
Ebihara A., K. Iwatsuki, T. A. Ohsawa and M. Ito. 2003. Hymenophyllum paniense (Hymenophyllaceae), a new species of filmy fern from New Caledonia. Syst. Bot. 28: 223-235.
Hennequin S., A. Ebihara, M. Ito, K. Iwatsuki and J.-Y. Dubuisson. 2003. Molecular systematics of the fern genus Hymenophyllum s.l. (Hymenophyllaceae) based on chloroplastic coding and noncoding regions. Molec. Phylogen. Evol. 27: 283-301.
Ebihara A., K. Iwatsuki, S. Kurita and M. Ito. 2002. Systematic position of Hymenophyllum rolandi-principis Rosenst. or a monotypic genus Rosenstockia Copel. (Hymenophyllaceae) endemic to New Caledonia. Acta Phytotax. Geobot. 53: 35-49.