Skeletonization 骨格化
In skeletonized leaves, only the veins and parts of the leaf margin remain, giving the leaf a “skeleton-like” appearance. This type of damage is commonly produced by both larvae and adults of leaf beetles (family Chrysomelidae). For example, the Japanese beetle (Popillia japonica) is well known for skeletonizing leaves, often leaving behind a network of veins. Certain chrysomelid larvae and some sawfly larvae (Tenthredinidae) can also produce skeletonized leaves during heavy feeding.

