A List of Pselaphine Species Known from BORNEO
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44 genera 123 spp.
Updated 31. Jul. 2008

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Literature(s):
Nomura, S. and M. Mohamed (2008) A faunistic review of the pselaphine species (Insecta, Coleoptera, Staphylinidae) known from borneo. Mem. natl. Mus. Nat. Sci., Tokyo, (45): 13-40.

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*In the checklist shown below, all valid generic names are given, with the number of species in parentheses, and the synonymic list is followed. In the synonymic list, each generic name is noted the original source and the status. Each genus is commented in Remarks on its systematic position and identity among similar taxa.
All valid scientific names and some important synonyms of pselaphine species known from Borneo are listed up. The valid specific names were coded as BN001, BN002, BN003 ...; BN is an abbreviation of �gBorneo�h. The important synonym(s) (indicated as Syn.) are listed after the valid name and coded as BN01A, B, C...<
For each scientific name of the genus and the species, author�fs name(s), year and the following symbols (a, b, c, �c) which are coincident with the list of literature shown at the tail, and the page number in original description is followed. The originally combined genus (if it was changed) is shown in parentheses.
The type locality is also added and distributional range of the species is indicated in square brackets.
** Goniacerine species described by Loebl et al., 2007 are not included.

Supertribe Bythinoplectitae
BN 001. Octomicrus divergens Reitter, 1884: 420. Borneo (Telang) [Borneo].
BN 002. Zethopsus simplicifrons Reitter, 1884: 419. Telang [Borneo].

Supertribe Euplectitae
BN 003. Aphilia femorata Reitter, 1883c: 296. Borneo (Tumbang-Hiang) [Borneo].

Supertribe Batrisitae
BN 004. Batrisodes? exiguus Raffray, 1882: 61. Java [Indonesia (Java, Sumatra, Borneo)].
BN 004A. Syn. B?. nicotianus Schaufuss, 1882b: 393. Sumatra.
BN 004B. Syn. B?. lateridens (Reitter, 1884: 398) (Batrisus). Telang, Tameanglaiang (Borneo).
BN 005. B?. incertus Schaufuss, 1882b: 392. Borneo, Sarawak [Borneo, New Guinea].
BN 006. Batrisoschema lateridentata Reitter, 1884: 400. Barabei [Borneo].
BN 007. Batrisus? grypochirus Schaufuss, 1882b: 379. Borneo, Sarawak [Borneo (Sarawak)].
BN 008. Borneana biformis Schaufuss, 1882c: 140. Borneo [Borneo].
BN 009. Mnia antennalis (Lobl, 1973: 191) (Mina). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 010. M. bella (Lobl, 1973: 194) (Mina). Sabah (Umas, near Tawau) [Borneo (Sabah)].
BN 011. M. brevicornis (Lobl, 1973: 164) (Mina). Sarawak (Semengoh, Mt. Matang) [Borneo (Sarawak)].
BN 012. M. brevipilis (Lobl, 1973: 169) (Mina). Sabah (Umas, near Tawau) [Borneo (Sabah)].
BN 013. M. bryanti (Lobl, 1973: 189) (Mina). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 014. M. fossulata (Lobl, 1973: 165) (Mina). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 015. M. frontalis (Lobl, 1973: 192) (Mina). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 016. M. gracilis (Lobl, 1973: 170) (Mina). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 017. M. macrops (Lobl, 1973: 175) (Mina). Sabah (Sibuga Forest Reserve) [Borneo (Sabah)].
BN 018. M. puncticeps (Lobl, 1973: 180) (Mina). Sarawak (Kampong Segu, near Kuching) [Borneo (Sarawak)].
BN 019. M. simulans (Lobl, 1973: 169) (Mina). Sarawak (Kampong Segu, near Kuching) [Borneo (Sarawak)].
BN 020. M. soror (Lobl, 1973: 168) (Mina). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 021. M. taylori (Lobl, 1973: 172) (Mina). Sarawak (Semengoh) [Borneo (Sarawak)].
BN 022. M. tuberculata (Lobl, 1973: 187) (Mina). Sarawak (Semengoh) [Borneo (Sarawak)]. BN 023. M. variabilis (Lobl, 1973: 178) (Mina). Sabah (Labuk Sepilok, near Sandakan) [Borneo (Sabah)].
BN 024. M. varians (Lobl, 1973: 168) (Mina). Sabah (Tawau) [Borneo (Sabah)].
BN 025. M. velaris (Lobl, 1973: 176) (Mina). Sabah (Tawau) [Borneo (Sabah)].
BN 026. Siteromina dayak Lobl, 1979: 191. Sarawak (Semengoh) [Borneo (Sarawak)].
BN 027. Sathytes punctiger Westwood, 1870: 128. Borneo (Sarawak) [Borneo (Sarawak)].
BN 028. Cratna pedestris Lobl, 1986: 223. Sarawak (Gunung Mulu) [Borneo (Sarawak)].
BN 029. Eubatrisus pubifer (Reitter, 1884: 397) (Batrisus (Batrisodes)). Borneo (Telang) [Borneo].
BN 030. Batrisocenus abbreviatus (Reitter, 1883: 285) (Batrisus (Batrisodes)). Borneo (Tumbang-Hiang) [Borneo].
BN 031. B. architectus (Reitter, 1884: 396) (Batrisus (Batrisodes)) Tameanglaiang [Borneo].
BN 032. B. batavianus (Reitter, 1883: 284) (Batrisus). Batavia [Borneo, Indonesia (Java)].
BN 033. B. bipunctulus (Reitter, 1884: 394) (Batrisus (Batrisodes)). Telang [Borneo].
BN 034. B. brevis (Schaufuss, 1882b: 396) (Batrisus). Borneo, Sarawak [Borneo].
BN 035. B. cavifer (Reitter, 1884: 395) (Batrisus (Batrisodes)). Telang [Borneo, Indonesia (Sumatra)].
BN 035A. Syn. Batrisus (Batrisodes) torticornis Raffray, 1892: 489. Bouchard (Sumatra).
BN 036. B. claviger (Reitter, 1884: 395) (Batrisus (Batrisodes)). Borneo (Telang) [Borneo].
BN 037. B. custos (Schaufuss, 1887: 138) (Batrisus). Borneo [Borneo].
BN 038. B. erectus (Raffray, 1896: 244) (Batrisodes). Borneo [Borneo].
BN 039. B. fundaebraccatus (Schaufuss, 1882a: 71) (Batrisus). Batavia (Sijthoff) [Singapore, Borneo (Telang Barabei), Indonesia (Java, Sumatra, Celebes), New Guinea].
BN 039A. Syn. Batrisus physoderes Schaufuss, 1882b: 383. Sumatra.
BN 039B. Syn. Batrisus nephriticus Schaufuss, 1882b: 384. Celebes, Macassar.
BN 039C. Syn. Batrisus similis Schaufuss, 1882b: 395. Sumatra.
BN 040. B. indus (Schaufuss, 1882b: 375) (Batrisus). Borneo, Sarawak [Borneo].
BN 041. B. laminidens (Reitter, 1884: 396) (Batrisus (Batrisodes)). Telang [Borneo].
BN 042. B. orbicollis (Reitter, 1884: 339) (Batrisus). Telang [Borneo].
BN 043. B. reitteri (Raffray, 1896: 249) (Batrisodes). Borneo [Borneo].
BN 044. B. sarawakensis (Schaufuss, 1882b: 381) (Batrisus). Borneo, Sarawak [Borneo].
BN 045. B. septemdentatus (Schaufuss, 1882b: 376) (Batrisus). Borneo, Sarawak [Borneo].
BN 046. B. spinidens (Reitter, 1884: 398) (Batrisus (Syrbatus)). Borneo (Telang) [Borneo].
BN 047. B. tarsalis (Reitter, 1884: 396) (Batrisus (Batrisodes)). Barabei, Telang [Borneo].
BN 048. B. vestigifer (Reitter, 1884: 394) (Batrisus (Batrisodes)). Telang, Tameanglaiang [Borneo].

Supertribe Goniaceritae
BN 049. Harmophorus parviceps (Reitter, 1884: 392) (Arnyllium). Barabei [Borneo].
BN 049A. Syn. Arnyllium longipile Raffray, 1892: 490 Sumatra.
BN 050. H. ensipes (Reitter, 1884: 392) (Arnyllium). Telang [Borneo].
BN 051. H. pectinatum (Reitter, 1884: 392) (Arnyllium). Telang, Tameanglaiang [Singapore, Borneo, Indonesia (Sumatra), Taiwan?].
BN 052. Comatopselaphus opacicollis Schaufuss, 1882b: 369. Borneo, Sarawak [Borneo]
BN 053. Reichenbachia affinissima Reitter, 1884: 402. Telang [Borneo].
BN 054. R. amica Reitter, 1884: 405. Tameanglaiang, Barabei [Borneo, Indonesia (Sumatra)].
BN 055. R. grabowskyi Reitter, 1883: 289. Borneo (Tumbang-Hiang) [Borneo].
BN 056. R. ingrata Reitter, 1884: 403. Telang [Borneo].
BN 057. R. integrostriata Reitter, 1884: 403. Telang [Borneo].
BN 058. R. invalida Reitter, 1883: 288. Borneo (Tumbang-Hiang) [Borneo].
BN 059. R. lamellicornis Reitter, 1883: 290. Borneo (Tumbang-Hiang) [Borneo].
BN 060. R. negligens Reitter, 1884: 403. Pengaron [Borneo].
BN 061. R. punctithorax Reitter, 1883: 290. Java (Batavia), Borneo (Tumbang-Hiang) [Indonesia (Java), Borneo]
BN 062. R. schaufussi Reitter, 1883: 289. Java (Batavia), Borneo (Tumbang-Hiang) [Singapore, Indonesia (Java, Sumatra), Borneo, Sri Lanka].
BN 062A. Syn. Reichenbachia extuscurvata Schaufuss, 1887: 114. Sumatra [Indonesia (Sumatra)].
BN 063. R. subvalida Reitter, 1884: 404. Telang [Borneo].
BN 064. R. telangensis Reitter, 1884: 402. Telang [Borneo].
BN 065. R. rufa (Schmidt-Gobel, 1838: 6) (Bryaxis). Ostindien [Singapore, India, Sri Lanka, Thailand, Vietnam, Borneo, Indonesia (Java, Celebes), Phillippines, S].
BN 066. Rybaxis amica Reitter, 1884: 405. Pengaron [Borneo].
BN 067. R. imperatrix (Schaufuss, 1882b: 363) (Bryaxis). Borneo, Sarawak [Borneo].
BN 068. R. nubila Reitter, 1883: 292. Borneo (Tumbang-Hiang) [Borneo].
BN 069. R. sarawakensis (Schaufuss, 1882b: 364) (Bryaxis). Borneo, Sarawak [Borneo].
BN 070. Eupines sphaerica (Motschulsky, 1851: 492) (Bryaxis). Indes or. [Singapore, Borneo, India, Sri Lanca, Myanmar, Indonesia (Sumatra, Java, Sulawesi), New Guinea, Japan (Ishigaki-jima Is., Iriomote-jima Is.)].
BN 082. Bythinophanax bicornis Reitter, 1884: 407. Telang [Borneo].
BN 083. B. exilis Reitter, 1884: 406. Tameanglaiang [Borneo].
BN 084. B. latebrosus Reitter, 1884: 406. Telang [Borneo].
BN 085. Bythinoderes grabowskyi Reitter, 1884: 408. Telang [Borneo].
BN 086. Brunomanseria faceta Lobl et Kurbatov, 2004: 366. Borneo (East Malaysia, Sabah, Crocket (Crocker) Range, km 51-52 Kota Kinabalu-Tanbunan, 1,600m) [Borneo]
BN 071. Plagiophorus baumeisteri (Reitter, 1884: 388) (Cyathiger). Telang, Tameanglaiang [Borneo].
BN 072. P. borneensis (Raffray, 1895: 55) (Cyathiger). Borneo [Borneo].
BN 073. P. bulbifer (Bryant, 1915b: 254) (Cyathiger). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 074. P. dispar (Bryant, 1915b: 251) (Cyathiger). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 075. P. elongatus (Bryant, 1915b: 252) (Cyathiger). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 076. P. matangensis (Bryant, 1915b: 253) (Cyathiger). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 077. P. mirandus (Bryant, 1915b: 250) (Cyathiger). Sarawak (Mt. Merinjak) [Borneo (Sarawak)].
BN 078. P. myrmecophilus (Bryant, 1915b: 250) (Cyathiger). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 079. P. sarawakensis (Bryant, 1915: 253) (Cyathiger). Sarawak (Quop) [Borneo (Sarawak)].
BN 080. P. schaufussi (Reitter, 1884: 385) (Cyathiger). Telang [Borneo].
BN 081. P. simonis (Reitter, 1884: 387) (Cyathiger). Telang, Tameanglaiang [Borneo].
BN 087. Atychodea lenticornis Reitter, 1884: 414. Telang, Tameanglaiang [Borneo].
BN 088. A. raffrayi Reitter, 1884: 414. Telang [Borneo].
BN 089. A. simoniana Reitter, 1884: 413. Telang [Borneo].
BN 090. A. singularis Reitter, 1884: 414. Tameanglaiang [Borneo].

Supertribe Pselaphitae
BN 091. Sabarhytus kinabalu Lobl, 2000: 147. Sabah (Mt. Kinabalu) [Borneo (Sabah)].
BN 092. Odontalgus vestitus Schaufuss, 1886: 243. Sumatra [Borneo?, Sumatra].
BN 093. Ctenistes mitis Schaufuss, 1882a: 74. Batavia (Sijthoff) [Vietnam (Saigon, Hiep Hoa), Borneo, Indonesia (Java, Sumatra)].
BN 094. Apharina borneensis Raffray, 1896: 270. Borneo [Borneo].
BN 095. A. fuscipennis Reitter, 1884: 415. Telang [Borneo].
BN 096. Hybocephalodes dentiventris (Reitter, 1884: 416) (Hybocephalus). Telang [Borneo].
BN 097. H. informis (Reitter, 1884: 417) (Hybocephalus). Telang, Tameanglaiang [Borneo].
BN 098. H. telangensis (Reitter, 1884: 417) (Hybocephalus). Telang, Tameanglaiang [Borneo].
BN 099. Pseudapharina prima (Reitter, 1884: 415) (Filiger). Telang [Borneo].
BN 100. Pselaphidius parvipalpis (Reitter, 1883: 294) (Pselaphus). Batavia [Borneo, Indonesia (Java, Sumatra), Singapore, Thailand, Vietnam (Saigon)].
BN 100A. Syn. Pselaphus bivestitus Schaufuss 1888: 295.
BN 101. Pselaphus unipunctatus Reitter, 1884: 410. Barabei, Pengaron [Borneo].
BN 102. P. lativentris Reitter, 1883: 293. Batavia [Borneo, Indonesia (Java)].
BN 104. P. calopygaeus Schaufuss, 1888: 294. Borneo; mer. or. [Borneo].
BN 105. P. canaliculatus Schaufuss, 1877: 5. Siam (Bangkok) [Thailand, Borneo (Sarawak)].
BN 106. P. sexstriatus Reitter, 1884: 411. Tameanglaiang [Borneo].
BN 107. P. biocellatus Reitter, 1884: 410. Telang, Tameanglaiang, Pengaron [Borneo].
BN 108. Chandleriella termitophila (Bryant, 1915c: 300) (Lasinus). Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 109. Tmesiphorus matangensis Bryant, 1915c: 297. Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 110. Centrophthalmus bispinus Reitter, 1884: 390. Telang [Borneo].
BN 111. C. divisus Reitter, 1884: 390. Telang [Borneo].
BN 112. C. punctipennis Schaufuss, 1877: 21. Siam (Bangkok) [Taiwan, Vietnam (Saigon), Thailand, Singapore, Borneo, Indonesia (Sumatra, Java, Sulawesi)].
BN 112A. Syn. C. punctatissimus Schaufuss, 1877: 22.
BN 112B. Syn. C. inaequalis Schaufuss, 1877: 22.
BN 112C. Syn. C. quadristriatus Schaufuss, 1877: 22. Singapore.
BN 113. Centrophthalmina laevis (Raffray, 1896: 276) (Centrophthalmus). Borneo [Vietnam (Saigon), Sumatra, Borneo].
BN 114. Enantius rostratus Reitter, 1884: 390. Tamaenglaiang [Borneo].
BN 115. Labomimus heterocerus (Raffray, 1882: 16) (Pselaphodes). Sarawak [Borneo (Sarawak); Java, Sumatra].
BN 115A. Syn. Pselaphodes sumatrensis Raffray, 1896: 277. Sumatra.
BN 115B. Syn. Pselaphodes antennatus Bryant, 1915c: 299. Sarawak (Quop).
BN 116. Pselaphodes villosus Westwood, 1870: 129. Borneo (Sarawak) [Borneo (Sarawak)].
BN 117. Tyrus myrmecophilus Bryant, 1915c: 298. Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 118. Stethotaphrus hamotoides (Schaufuss, 1882b): 370 (Taphrostethus). Borneo, Sarawak [Borneo].

Supertribe Clavigeritae
BN 119. Colilodion incredibilis Besuchet, 1991: 503. Sabah (Mt. Kinabalu) [Borneo (Sabah)].
BN 120. C. inopinatus Besuchet, 1991: 507. Sabah (Tamparuli-Ranau) [Borneo (Sabah)].
BN 121. C. mirus Besuchet, 1991: 506. Sabah (Mt. Kinabalu) [Borneo (Sabah)].
BN 122. Articerodes borneensis Bryant, 1915a: 213. Sarawak (Mt. Matang) [Borneo (Sarawak)].
BN 123. Disarthricerus moultoni Bryant, 1915 a: 212. Sarawak (Mt. Matang) [Borneo (Sarawak)].

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